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Two archosaur branches, eleven evidence dossiers

Keep named specimens, feather chemistry, wind-tunnel measurements, phylogenetic topology, molecular clocks and COL26.8 living-name coverage separate while comparing crocodile-line and bird-line evidence.

Each step names its evidence claims, with references below. Interactive Explorer states are available in the full application; the narrative remains an editorial synthesis.

Story sequence

01 · 205 Ma · evidence

A suchian can resemble a theropod

AMNH FR 30587 records an edentulous, gracile suchian. Its ornithomimid-like proportions are a convergence diagnosis, not evidence that the two branches share an immediate ancestor or identical ecology.

Effigia is a pseudosuchian outside Crocodylomorpha, so it remains event evidence rather than a false child of the crocodile navigation node.

claim:event:effigia-convergent-body-plan
02 · 231 Ma · evidence

A partial holotype, a bounded predator inference

NCSM 21558 documents a large Carnian crocodylomorph. Size and predator tier come from proxies and assemblage context, not preserved feeding behaviour.

Incomplete and immature anatomy cannot establish a direct ancestor or universal global first occurrence.

claim:event:carnufex-holotype-predator
03 · 165.9 Ma · evidence

Skull consolidation appears as a mosaic

IVPP V14010 combines a consolidated, crocodylian-like skull with a gracile terrestrial postcranium. It is one anatomical mosaic rather than a compulsory stage in a ladder.

Skull mechanics and cursoriality are functional interpretations; topology is character-matrix evidence.

claim:event:junggarsuchus-skull-consolidation
04 · 0 Ma · tree

Living gharials meet fossils in a tip-dated model

A combined morphology, molecular and age model groups Gavialis with Tomistoma while moving sampled thoracosaurs outside the crown. Other analyses differ.

A preferred topology is not an observed genealogy; priors, weights and sampled characters remain visible boundaries.

claim:event:crocodylian-tip-dated-topology
05 · 149 Ma · evidence

Archaeopteryx preserves an anatomical mosaic

WDC CSG 100 exposes theropod-like pedal and palate features alongside avialan anatomy. The mosaic supports affinity without turning one specimen into the unique first bird.

Joint posture, perching and exact placement are interpretations rather than direct observations.

claim:event:archaeopteryx-thermopolis-skeleton
06 · 160 Ma · evidence

Microscopy informs a colour reconstruction

Melanosomes in BMNHC PH828 support a statistical map of plumage regions. Preservation, reference samples and a tail borrowed from another specimen constrain the image.

Anchiornis remains a non-avian paravian event here, not a false child of Aves or a directly observed ancestral colour state.

claim:event:anchiornis-plumage-melanosomes
07 · 122.5 Ma · evidence

Four feathered surfaces are preserved

IVPP V13352 preserves long pennaceous feathers on forelimbs, hindlimbs and tail. The fossil records surfaces, not a flight trajectory.

Microraptor is treated as a non-avian paravian dossier, not a direct bird ancestor.

claim:event:microraptor-four-winged-holotype
08 · 0 Ma · evidence

A wind tunnel tests feasibility

Measured forces on a scaled model support stable high-drag gliding across tested configurations. That experiment is functional evidence, not footage of extinct behaviour.

Geometry, scaling, reconstructed posture and fossil distortion bound every aerodynamic conclusion.

claim:event:microraptor-wind-tunnel-model
09 · 66.75 Ma · tree

One skull, two nearby crown placements

CT anatomy anchors NHMM 2013 008 near Galloanserae, but parsimony and tip-dated Bayesian analyses place it on different branches.

A tightly dated occurrence is not automatically the globally oldest modern bird, and crown membership remains analysis-dependent.

claim:event:asteriornis-holotype-crown-placement
10 · 0 Ma · tree

Whole genomes still retain difficult branches

Forty-eight representatives across neoavian orders recover major groupings and expose gene-tree conflict. This is sampled extant topology, not a complete species tree.

Incomplete lineage sorting, convergence, sampling and method choice remain explicit.

claim:event:neoavian-genome-topology
11 · 0 Ma · tree

A calibrated time tree is a model

Clock-like exons, broader RAG sampling and fossils infer a Cretaceous root and boundary-era radiation. The dates follow calibrations, topology and priors.

COL26.8 routes 11,071 accepted living species into this package. That is a nomenclatural coverage boundary, not fossil completeness, direct ancestry or agreement on a single crown-bird date.

claim:event:neornithes-fossil-calibrated-time-tree

References

  1. Extreme convergence in the body plans of an early suchian (Archosauria) and ornithomimid dinosaurs (Theropoda)Nesbitt, S.J.; Norell, M.A. · 2006 · DOI 10.1098/rspb.2005.3426
  2. Early crocodylomorph increases top tier predator diversity during rise of dinosaursZanno, L.E.; Drymala, S.; Nesbitt, S.J.; Schneider, V.P. · 2015 · DOI 10.1038/srep09276
  3. A Middle Jurassic 'sphenosuchian' from China and the origin of the crocodylian skullClark, J.M.; Xu, X.; Forster, C.A.; Wang, Y. · 2004 · DOI 10.1038/nature02802
  4. Tip-dating and homoplasy: reconciling the shallow molecular divergences of modern gharials with their long fossil recordLee, M.S.Y.; Yates, A.M. · 2018 · DOI 10.1098/rspb.2018.1071
  5. A well-preserved Archaeopteryx specimen with theropod featuresMayr, G.; Pohl, B.; Peters, D.S. · 2005 · DOI 10.1126/science.1120331
  6. Plumage color patterns of an extinct dinosaurLi, Q.; Gao, K.-Q.; Vinther, J.; et al. · 2010 · DOI 10.1126/science.1186290
  7. Four-winged dinosaurs from ChinaXu, X.; Zhou, Z.; Wang, X.; Kuang, X.; Zhang, F.; Du, X. · 2003 · DOI 10.1038/nature01342
  8. Aerodynamic performance of the feathered dinosaur Microraptor and the evolution of feathered flightDyke, G.; de Kat, R.; Palmer, C.; van der Kindere, J.; Naish, D.; Ganapathisubramani, B. · 2013 · DOI 10.1038/ncomms3489
  9. Late Cretaceous neornithine from Europe illuminates the origins of crown birdsField, D.J.; Benito, J.; Chen, A.; Jagt, J.W.M.; Ksepka, D.T. · 2020 · DOI 10.1038/s41586-020-2096-0
  10. Whole-genome analyses resolve early branches in the tree of life of modern birdsJarvis, E.D.; Mirarab, S.; Aberer, A.J.; et al. · 2014 · DOI 10.1126/science.1253451
  11. A new time tree reveals Earth history's imprint on the evolution of modern birdsClaramunt, S.; Cracraft, J. · 2015 · DOI 10.1126/sciadv.1501005