Stories / 9 minHow early land-plant evidence changes shape
Move from a modelled crown divergence to dispersed cryptospores, spore-bearing fragments and anatomically informative Devonian bodies without collapsing them into one origin date.
Each step names its evidence claims, with references below. Interactive Explorer states are available in the full application; the narrative remains an editorial synthesis.
Story sequence
01 · 494 Ma · evidenceA modelled crown interval
A fossil-calibrated relaxed clock places crown Embryophyta in a broad middle Cambrian–Early Ordovician interval, but this is a model result rather than a fossil occurrence.
The Plantae node is navigation context; the claim concerns crown Embryophyta.
claim:event:plants-on-land02 · 472 Ma · mapDispersed spores on emerged land
A Dapingian assemblage in Argentina contains five cryptospore genera, but dispersed propagules do not reveal a complete parent plant or a crown-bryophyte identity.
The source-reported numerical ages are retained; they are not silently replaced by the current ICS Dapingian boundaries.
claim:event:dapingian-cryptospores03 · 451 Ma · evidenceSpores linked to plant fragments
Late Ordovician Oman fragments preserve dense spore masses and probable sporangial covering, strengthening the land-plant interpretation while leaving liverwort crown affinity tentative.
The broad window translates a legacy Caradoc assignment for display; it is not a point date.
claim:event:ordovician-sporangia:caradoc-record · claim:event:ordovician-sporangia:spore-masses04 · 407 Ma · treeA three-axis rooting system
Three-dimensional fossil reconstruction shows how Asteroxylon combined leafy shoots, root-bearing axes and rooting axes through an extinct developmental pathway.
A species-level exemplar cannot supply morphology for every lycophyte lineage.
claim:event:asteroxylon-rooting-system05 · 385 Ma · evidenceA liverwort body-fossil record
Givetian Metzgeriothallus preserves thalloid gametophytes and an associated sporophyte capsule, adding body-plan information without turning one occurrence into a phylum origin date.
Occurrence, morphology and crown divergence remain separate evidence objects.
claim:event:metzgeriothallus:givetian-record · claim:event:metzgeriothallus:body-planReferences
- The timescale of early land plant evolutionMorris, J.L.; Puttick, M.N.; Clark, J.W.; et al. · 2018 · DOI 10.1073/pnas.1719588115
- Early Middle Ordovician evidence for land plants in Argentina (eastern Gondwana)Rubinstein, C.V.; Gerrienne, P.; de la Puente, G.S.; Astini, R.A.; Steemans, P. · 2010 · DOI 10.1111/j.1469-8137.2010.03433.x
- Fragments of the earliest land plantsWellman, C.H.; Osterloff, P.L.; Mohiuddin, U. · 2003 · DOI 10.1038/nature01884
- An evidence-based 3D reconstruction of Asteroxylon mackiei, the most complex plant preserved from the Rhynie chertHetherington, A.J.; Bridson, S.L.; Jones, A.L.; Hass, H.; Kerp, H.; Dolan, L. · 2021 · DOI 10.7554/eLife.69447
- Earth's oldest liverworts—Metzgeriothallus sharonae sp. nov. from the Middle Devonian (Givetian) of eastern New York, USAVanAller Hernick, L.; Landing, E.; Bartowski, K.E. · 2008 · DOI 10.1016/j.revpalbo.2007.09.002