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故事 / 7 分钟

从立体网骨片到现生棘皮动物冠群

Follow five evidence boundaries that keep isolated skeleton, contested affinity, articulated bodies and modelled crown time distinct.

每一步标明所依据的主张,引用列于下方。交互探索状态可在完整应用中打开;故事本身仍是编辑综合。

故事步骤

01 · 522 Ma · 证据

An ossicle is not an articulated animal

Cambrian Stage 3 stereom ossicles directly record echinoderm skeletal microstructure, but isolated elements do not reveal one complete body plan or a phylum origin date.

The source-reported numerical calibration is approximate and older than the atlas's current chart.

claim:taxon:echinodermata:earliest-stereom-ossicles
02 · 536 Ma · 树

Keep a Fortunian candidate contested

Yanjiahella was recovered as a stem echinoderm in one analysis, but a published reanalysis found no unambiguous echinoderm synapomorphy or stereom, so it does not set the range.

Competing matrices are evidence of disagreement, not permission to select a convenient endpoint.

claim:taxon:echinodermata:yanjiahella-affinity
03 · 510 Ma · 多样性

Articulated faunas change the record

By about 510 Ma, articulated echinoderms preserve at least four markedly differentiated body plans, supplying a more robust body-fossil datum than isolated stereom.

A robust sampled fauna is still not the first evolutionary instant of Echinodermata.

claim:taxon:echinodermata:articulated-faunas-510ma
04 · 475 Ma · 地图

Class-level records remain uneven

The unproblematic crinoid record begins in the middle Tremadocian, while the oldest articulated holothurian used here is late Darriwilian; each class keeps its own specimen and stratigraphic boundary.

Different preservation histories prevent one class record from standing in for every echinoderm class.

claim:taxon:crinoidea:tremadocian-record · claim:taxon:holothuroidea:darriwilian-articulated-record
05 · 465 Ma · 树

Do not merge stem fossils with crown time

Darriwilian bothriocidaroid plates anchor total-group Echinoidea, whereas phylogenomic analyses place the crown near the Carboniferous–Permian transition under model-dependent clocks.

A fossil occurrence and a modelled divergence answer different questions and remain separate in the atlas.

claim:taxon:echinoidea:ordovician-stem-record · claim:taxon:echinoidea:crown-origin

参考文献

  1. The oldest echinoderm faunas from Gondwana show that echinoderm body plan diversification was rapidSmith, A.B.; Zamora, S.; Álvaro, J.J. · 2013 · DOI 10.1038/ncomms2391
  2. A stem group echinoderm from the basal Cambrian of China and the origins of AmbulacrariaTopper, T.P.; Guo, J.; Clausen, S.; Skovsted, C.B.; Zhang, Z. · 2019 · DOI 10.1038/s41467-019-09059-3
  3. Re-evaluating the phylogenetic position of the enigmatic early Cambrian deuterostome YanjiahellaZamora, S.; Wright, D.F.; Mooi, R.; Lefebvre, B.; Guensburg, T.E.; et al. · 2020 · DOI 10.1038/s41467-020-14920-x
  4. Athenacrinus n. gen. and other early echinoderm taxa inform crinoid origin and arm evolutionGuensburg, T.E.; Sprinkle, J.; Mooi, R.; Lefebvre, B.; David, B.; Roux, M.; Derstler, K.L. · 2020 · DOI 10.1017/jpa.2019.87
  5. Fauna and ecology of the holothurian bed, Llandrindod, Wales, UK (Darriwilian, Middle Ordovician), and the oldest articulated holothurianBotting, J.P.; Muir, L.A. · 2012 · DOI 10.26879/272
  6. The Paleobiology Database application programming interfacePeters, S.E.; McClennen, M. · 2016 · DOI 10.1017/pab.2015.39
  7. The Ordovician diversification of sea urchins: systematics of the Bothriocidaroida (Echinodermata: Echinoidea)Thompson, J.R.; Cotton, L.J.; Candela, Y.; Kutscher, M.; Reich, M.; Bottjer, D.J. · 2022 · DOI 10.1080/14772019.2022.2042408
  8. Phylogenomic analyses of echinoid diversification prompt a re-evaluation of their fossil recordMongiardino Koch, N.; Thompson, J.R.; Hiley, A.S.; McCowin, M.F.; Armstrong, A.F.; et al. · 2022 · DOI 10.7554/eLife.72460