Named Burgess Shale specimens expose mandibles, maxillipeds and basipod endites used to test the early mandibulate body plan.
Represented range508–508 Ma
RegionsMarble Canyon, Burgess Shale, British Columbia, Canada
CladesMandibulata · Tokummia katalepsis
Claim ledger
morphology · high
Holotype ROM 63823 and paratypes of Tokummia katalepsis preserve paired mandibles, maxillipeds and subdivided basipods; their recovery as stem mandibulates is a morphology-matrix result, not evidence that Tokummia is a direct ancestor of crustaceans, insects or myriapods.