References / paperA marine stem-myriapod from the Silurian Waukesha Lagerstätte, Wisconsin, USA: terrestrial traits pre-date the transition to land
Briggs, D.E.G.; Lamsdell, J.C.; Kluessendorf, J.; Mikulic, D.G.
Year2026
Source roleprimary-study
DOI / ID10.1098/rspb.2026.0131
Fitness and scope
taxonomy · topology · range · morphology
Thirty-five Llandoverian Waukartus specimens preserve uniramous limbs, muscles and endoskeleton; morphology matrices place the taxon on the myriapod stem rather than within crown Myriapoda.
Claim ledger
fossil-range · mediumThe Myriapoda root display replaces the legacy 443 Ma value with a 437–0 Ma total-group navigation envelope anchored by Llandoverian Waukartus and living myriapods; Waukartus is recovered on the stem and is not a crown first appearance.
claim:taxon:myriapoda:root-range-rc39