References / paperThe earliest known holometabolous insects
Nel, A.; Roques, P.; Nel, P.; Prokin, A.A.; Bourgoin, T.; Prokop, J.; et al.
Year2013
Source roleprimary-study
DOI / ID10.1038/nature12629
Fitness and scope
taxonomy · range · morphology · topology
Describes five Pennsylvanian fossils representing a stem coleopterid, a holometabolous larva, a stem hymenopterid, Hemiptera and Psocodea.
Claim ledger
fossil-range · mediumFive Moscovian-to-Gzhelian insect fossils document a stem coleopterid, a holometabolous larva, a stem hymenopterid, early Hemiptera and Psocodea, expanding sampled Pennsylvanian Eumetabola without proving synchronous origins or later crown-order first appearances.
claim:event:pennsylvanian-eumetabola-sampletopology · mediumThe Insect radiation evidence route navigation entity is source-linked only to the named specimen, dataset and analysis dossiers already cited by its range ledger. Its displayed span is a study-bounded route or sample envelope, not a biological taxon, ancestor sequence, global FAD/LAD or exact origin interval.
claim:taxon:insect-radiation-evidence-routefossil-range · mediumPennsylvanian Eumetabola sample is bounded here by five fossils from Moscovian and Gzhelian localities in France and Russia; this composite sample is not one lineage range or a complete Eumetabola history.
claim:taxon:pennsylvanian-eumetabola