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Other mammals: twelve evidence dossiers, no ancestor ladder

Compare named fossils, a living genome, morphology matrices, bone-density measurements and molecular trees while keeping specimen facts, function, topology, clocks and COL26.8 name routing separate.

Each step names its evidence claims, with references below. Interactive Explorer states are available in the full application; the narrative remains an editorial synthesis.

Story sequence

01 · 105 Ma · evidence

Steropodon jaw and monotreme dental evidence

AM F66763 preserves a right dentary fragment and three molars whose morphology was compared with monotreme and tribosphenic conditions.

Dental homology and affinity are interpretations; one jaw cannot establish a direct ancestor, a complete body plan or an immutable global first occurrence.

claim:event:steropodon-holotype-monotreme
02 · 0.0005 Ma · tree

Platypus draft genome and comparative mosaic

Approximately 26.9 million reads produced a 1.84-gigabase draft at about sixfold coverage, with 437 megabases ordered on 20 chromosomes and 18,527 predicted protein-coding genes.

Assembly gaps, one-individual sampling, annotation and comparative models prevent direct reconstruction of an ancestral genome or single-gene explanations for organismal traits.

claim:event:platypus-genome-mosaic
03 · 124.94999999999999 Ma · diversity

Eomaia skeleton, topology and locomotor inference

CAGS 01-IG-1a,b documents the skull, dentition and much of the postcranium; the original matrix and limb comparisons supported a basal eutherian and scansorial interpretation.

Later combined matrices can move Eomaia outside Theria; topology and scansoriality are analytical outputs, not a directly observed ancestor or behaviour.

claim:event:eomaia-holotype-topology
04 · 160 Ma · evidence

Juramaia specimen and conditional Jurassic signal

BMNH PM1343B preserves skull, dentition and postcranium; the original study attributed it to approximately 160-million-year-old strata and recovered a basal eutherian position.

Commercial collection provenance, stratigraphic attribution and matrices condition the result; unconstrained tip dating yields a younger morphological age signal rather than directly redating the rock.

claim:event:juramaia-conditional-provenance
05 · 125.5 Ma · tree

Ambolestes and a revised therian boundary

STM33-5 preserves ectotympanic, malleus and hyoid anatomy; the published matrix recovers Ambolestes and Sinodelphys in Eutheria.

Reassignment and the resulting metatherian ghost lineage depend on sampled characters and taxa; the analysis does not observe the placental–marsupial split.

claim:event:ambolestes-therian-boundary
06 · 65 Ma · diversity

Placental phenomic matrix and K–Pg model

The study scored 4,541 phenomic characters for 86 living and fossil species, supported by more than 12,000 annotated images, and combined them with 27 nuclear genes.

The post-K–Pg chronology and reconstructed phenotype depend on parsimony, fossil ages, ghost lineages and ancestral-state optimization; no ancestor was directly observed.

claim:event:placental-phenomic-kpg-model
07 · 91 Ma · evidence

Placental four-clade molecular topology

The 16,397-base-pair matrix combines 19 nuclear and 3 mitochondrial genes; Bayesian and maximum-likelihood analyses recover four major placental clades.

The Afrotheria root, dates and Gondwanan scenario are model-conditioned, later studies test alternatives, and sampled taxa from other packages remain cross-boundary inputs.

claim:event:placental-molecular-four-clades
08 · 59.85 Ma · tree

Eritherium skull and stem-proboscidean placement

Holotype MNHN PM69 preserves a rostrum and maxillary dentition; fourteen additional specimens extend the sampled upper and lower teeth and skull anatomy.

Stem placement and estimated 3–8 kilogram body mass derive from a character matrix and dental allometry, not observed ancestry or body mass.

claim:event:eritherium-holotype-proboscidean
09 · 5.5 Ma · diversity

Thalassocnus bone-density series

CT slices and thin sections show increasing compactness across ribs and limb bones, including named T. littoralis specimens MNHN.F.SAS 2 and MNHN.F.SAS 53.

Aquatic adaptation, hydrostatic control and marine grazing are functional interpretations; species succession is not a directly observed ancestor chain.

claim:event:thalassocnus-bone-density-series
10 · 44.5 Ma · evidence

Mimolagus teeth, tarsals and body-size model

Holotype right M3 IVPP V20115 and referred teeth, astragalus, calcanei and cuboid establish the anatomical sample and locality.

A 4.5-kilogram estimate, cursoriality and tapiroid-like niche are regression and ecomorphology inferences; the taxon is not a direct lagomorph ancestor.

claim:event:mimolagus-holotype-glires
11 · 51.4 Ma · tree

Onychonycteris flight and echolocation proxies

ROM 55351A preserves elongated wings and a relatively small cochlea; limb proportions and cochlear size were compared with living and fossil bats.

Powered flight and absence of specialized laryngeal echolocation are functional inferences, not observed behaviour, and one specimen is not a direct ancestor.

claim:event:onychonycteris-flight-echolocation
12 · 0.0005 Ma · diversity

Seven-gene bat topology and echolocation scenarios

Four nuclear and three mitochondrial genes contributed 8,230 aligned base pairs; analyses group sampled rhinolophoids with megabats rather than other microbats.

Multiple origins or losses of laryngeal echolocation are alternative historical reconstructions; the topology does not directly observe the first flight or sonar event. COL26.8 routes 5,099 accepted living species to other-mammals after excluding Cetartiodactyla, Perissodactyla, Primates and Carnivora; that nomenclatural snapshot is not 5,099 dossiers and does not resolve fossil topology.

claim:event:bat-seven-gene-topology

References

  1. First Mesozoic mammal from Australia—an Early Cretaceous monotremeArcher, M.; Flannery, T.F.; Ritchie, A.; Molnar, R.E. · 1985 · DOI 10.1038/318363a0
  2. Genome analysis of the platypus reveals unique signatures of evolutionWarren, W.C. et al. · 2008 · DOI 10.1038/nature06936
  3. The earliest known eutherian mammalJi, Q.; Luo, Z.-X.; Yuan, C.-X.; Wible, J.R.; Zhang, J.-P.; Georgi, J.A. · 2002 · DOI 10.1038/416816a
  4. The placental mammal ancestor and the post–K–Pg radiation of placentalsO'Leary, M.A. et al. · 2013 · DOI 10.1126/science.1229237
  5. A Jurassic eutherian mammal and divergence of marsupials and placentalsLuo, Z.-X.; Yuan, C.-X.; Meng, Q.-J.; Ji, Q. · 2011 · DOI 10.1038/nature10291
  6. Tip dating supports novel resolutions of controversial relationships among early mammalsCelik, M.A.; Phillips, M.J. · 2020 · DOI 10.1098/rspb.2020.0943
  7. An Early Cretaceous eutherian and the placental–marsupial dichotomyBi, S.; Zheng, X.; Wang, X.; Cignetti, N.E.; Yang, S.; Wible, J.R. · 2018 · DOI 10.1038/s41586-018-0210-3
  8. Resolution of the early placental mammal radiation using Bayesian phylogeneticsMurphy, W.J. et al. · 2001 · DOI 10.1126/science.1067179
  9. Paleocene emergence of elephant relatives and the rapid radiation of African ungulatesGheerbrant, E. · 2009 · DOI 10.1073/pnas.0900251106
  10. Gradual adaptation of bone structure to aquatic lifestyle in extinct sloths from PeruAmson, E.; de Muizon, C.; Laurin, M.; Argot, C.; de Buffrénil, V. · 2014 · DOI 10.1098/rspb.2014.0192
  11. A large mimotonid from the Middle Eocene of China sheds light on the evolution of lagomorphs and their kinFostowicz-Frelik, Ł.; Li, C.; Mao, F.; Meng, J.; Wang, Y. · 2015 · DOI 10.1038/srep09394
  12. Primitive Early Eocene bat from Wyoming and the evolution of flight and echolocationSimmons, N.B.; Seymour, K.L.; Habersetzer, J.; Gunnell, G.F. · 2008 · DOI 10.1038/nature06549
  13. Molecular evidence regarding the origin of echolocation and flight in batsTeeling, E.C.; Scally, M.; Kao, D.J.; Romagnoli, M.L.; Springer, M.S.; Stanhope, M.J. · 2000 · DOI 10.1038/35003188