References / paperThe placental mammal ancestor and the post–K–Pg radiation of placentals
O'Leary, M.A. et al.
Year2013
Source roleprimary-study
DOI / ID10.1126/science.1229237
Fitness and scope
topology · range · morphology · geochronology
Primary combined-data analysis using MorphoBank phenomic characters and GenBank sequences; reconstructed anatomy and node ages are model outputs.
Claim ledger
topology · mediumCAGS 01-IG-1a,b preserves an articulated skeleton with dentition, feet and claws; the original matrix placed Eomaia near the eutherian root and inferred scansoriality, while later combined matrices can place it outside Theria, so neither topology nor locomotor habit is a directly observed ancestral state.
claim:event:eomaia-holotype-topologydivergence-time · mediumA matrix of 4,541 phenomic characters for 86 living and fossil species, combined with 27 nuclear genes, produced one parsimony tree with post-K–Pg crown-placental diversification and a reconstructed ancestral phenotype; both chronology and phenotype are model outputs rather than observations of an ancestor.
claim:event:placental-phenomic-kpg-modeltaxonomy · contestedCAGS 01-IG-1a,b was recovered as basal Eutheria in the original matrix, while later combined analyses can move Eomaia outside Theria; neither result is ancestry.
claim:taxon:eomaia:taxonomybiogeography · mediumThe profile is restricted to the Yixian Formation occurrence in Liaoning and does not establish a global eutherian or therian origin area.
claim:taxon:eomaia:biogeographyecology · mediumDentition and limb proportions support faunivorous and scansorial interpretations, while feeding and locomotor behaviour were not directly observed.
claim:taxon:eomaia:ecologymorphology · mediumThe articulated holotype preserves skull, dentition, much of the postcranium, feet and claws used in anatomical and phylogenetic comparisons.
claim:taxon:eomaia:morphologyfossil-range · mediumThe 125.8–124.1 Ma envelope is the Yixian sample context, not a universal therian FAD, eutherian calibration or direct specimen date.
claim:taxon:eomaia:fossil-rangefossil-range · contestedThe 100–0 Ma Placentalia route is a model envelope: a molecular analysis estimated a roughly 103 Ma deep placental split, whereas the phenomic combined analysis placed sampled crown diversification after K–Pg; neither model is a fossil FAD or ancestor observation.
claim:taxon:placentalia:root-rangefossil-range · lowThe 58.3–0 Ma Xenarthra route retains the Riostegotherium entry from a combined phenomic–molecular chronology as a sampled/model bound; it is not a securely dated global FAD, crown origin or complete fossil range.
claim:taxon:xenarthra:root-rangefossil-range · lowThe 66–0 Ma Glires route is a K–Pg-to-present navigation/model envelope contextualized by post-K–Pg placental analyses and a 47.8–41.2 Ma Mimolagus fossil sample; it does not date crown Glires or assert continuous fossil occupancy.
claim:taxon:glires:root-range